Initial Statement of Beneficial Ownership

This filing reports an initial statement of beneficial ownership concerning Series W-7 Variable Rate Muni Term Preferred Shares of BLACKROCK MUNIHOLDINGS FUND, INC. (MHD). The shares were beneficially owned by DNT Asset Trust and subsequently deposited into a trust where DNT Trust retains indirect beneficial ownership. JPMorgan Chase & Co. also holds an indirect interest through its subsidiary, DNT Trust. Both entities are reporting this as a 10% owner transaction. The transaction date was June 2, 2026, with the filing date of June 4, 2026. This filing is made jointly by JPMorgan Chase & Co. and DNT Asset Trust.