Form 4 Filing for Esther Martinborough - CODEXIS, INC.

This filing details a Form 4 submission by Esther Martinborough, a director at CODEXIS, INC. (CDXS). On June 17, 2026, Ms. Martinborough acquired 66,350 stock options with a right to buy at a price of $0.00 per share. These options are exercisable and pertain to the company's Common Stock. The options vest on the earlier of the first anniversary of the grant date or the next annual stockholder meeting, contingent upon continued service. The post-transaction shares owned following this acquisition are 66,350. The filing was signed on June 18, 2026, by Georgia Erbez as Attorney-in-Fact for Esther Martinborough.